Ursidae is a family of mammals in the order Carnivora, which includes the giant panda, brown bear, and polar bear, and many other extant and extinct mammals. A member of this family is called a bear or an ursid. They are widespread across the Americas and Eurasia. Bear habitats are generally forests, though some species can be found in grassland and savana regions, and the polar bear lives in arctic and aquatic habitats. Most bears are 1.2–2m (4–7ft) long, plus a 3–20cm (1–8in) tail, though the polar bear is 2.2–2.44m (7–8ft) long, and some subspecies of brown bear can be up to 2.8m (9ft). Weights range greatly from the sun bear, which can be as low as 35kg (77lb), to the polar bear, which can be as high as 726kg (1,600lb). Population sizes vary, with six species classified as vulnerable with populations as low as 500, while the brown bear has a population of over 100,000 and the American black bear around 800,000. Many bear species primarily eat specific foods, such as seals for the polar bear or termites and fruit for the sloth bear, but with the exception of the giant panda, which exclusively eats bamboo, ursids are omnivorous when necessary. No ursid species have been domesticated, though some bears have been trained for entertainment.[1]
Species in mammal family Ursidae
Brown bear (Ursus arctos)
The eight species of Ursidae are split into five genera in three subfamilies: the monotypic Ailuropodinae, the panda bears; Tremarctinae, the short-faced bears; and Ursinae, containing all other extant bears. Extinct species have also been placed into all three extant subfamilies, as well as three extinct ones: Agriotheriinae, Hemicyoninae, and Ursavinae. Over 100 extinct Ursidae species have been found, though due to ongoing research and discoveries the exact number and categorization is not fixed.
Conventions
IUCN Red List categories
Conservation status
EX
Extinct (0 species)
EW
Extinct in the wild (0 species)
CR
Critically Endangered (0 species)
EN
Endangered (0 species)
VU
Vulnerable (6 species)
NT
Near threatened (0 species)
LC
Least concern (2 species)
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "". Population figures are rounded to the nearest hundred.
Classification
The family Ursidae consists of eight extant species belonging to five genera in three subfamilies and divided into dozens of extant subspecies. This does not include ursid hybrid species such as grizzly–polar bear hybrids or extinct prehistoric species.
Subfamily Ailuropodinae
Genus Ailuropoda (panda bears): one species
Subfamily Tremarctinae
Genus Tremarctos (short-faced bears): one species
Subfamily Ursinae
Genus Helarctos: (sun bear): one species
Genus Melursus: (sloth bear): one species
Genus Ursus (bears): four species
Ursidae
Ursinae
Ursus (American black / Asian black / brown / polar bear)
Helarctos (sun bear)
Melursus (sloth bear)
Tremarctinae
Tremarctos (spectacled bear)
Ailuropodinae
Ailuropoda (giant panda)
Ursids
The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis; this includes the division of the giant panda into two subspecies. There are several additional proposals which are disputed, such as reclassifying the subspecies of the brown bear into a smaller set of clades,[2][3] which are not included here.
Subfamily Ailuropodinae
Main article: Ailuropodinae
Genus Ailuropoda (H. Milne-Edwards, 1870) – one species
In addition to extant bears, a number of prehistoric species have been discovered and classified as a part of Ursidae. In addition to being placed within the three extant subfamilies, they have been categorized within the extinct subfamilies Agriotheriinae, Hemicyoninae, and Ursavinae, some of which are subdivided into named tribes. There is no generally accepted classification of extinct ursid species. The species listed here are based on data from the Paleobiology Database, unless otherwise cited. Where available, the approximate time period the species was extant is given in millions of years before the present (Mya), also based on data from the Paleobiology Database.[26] All listed species are extinct; where a genus or subfamily within Ursidae comprises only extinct species, it is indicated with a dagger symbol .
Extinct prehistoric subspecies of an extant species
References
Findeizen, Nikolai (2008). History of Music in Russia from Antiquity to 1800, Vol. 1: From Antiquity to the Beginning of the Eighteenth Century. Indiana University Press. p.201. ISBN978-0-253-02637-8.
Calvignac, S.; Hughes, S.; Tougard, C.; Michaux, J.; Thevenot, M.; Philippe, M.; Hamdine, W.; Hanni, C. (2008). "Ancient DNA evidence for the loss of a highly divergent brown bear clade during historical times". Molecular Ecology. 17 (8): 1962–1970. doi:10.1111/j.1365-294x.2008.03631.x. PMID18363668. S2CID23361337.
Bies, LeeAnn (2002). "Ailuropoda melanoleuca". Animal Diversity Web. University of Michigan. Retrieved March 24, 2020.
"Physical Description". Knowledge Hub. World Wide Fund for Nature. Retrieved March 24, 2020.
Swaisgood, R.; Wang, D.; Wei, F. (2017) [errata version of 2016 assessment]. "Ailuropoda melanoleuca". IUCN Red List of Threatened Species. 2016: e.T712A121745669.
Ramesh, T.; Kalle, R.; Sankar, K.; Qureshi, Q. (2012). "Factors affecting habitat patch use by sloth bears in Mudumalai Tiger Reserve, Western Ghats, India". Ursus. 23 (1): 78–85. doi:10.2192/URSUS-D-11-00006.1. JSTOR41818973. S2CID86410436.
Dewey, Tanya; Kronk, Christine (2007). "Ursus americanus". Animal Diversity Web. University of Michigan. Retrieved March 24, 2020.
Garshelis, D. L.; Scheick, B. K.; Doan-Crider, D. L.; Beecham, J. J.; Obbard, M. E. (2017) [errata version of 2016 assessment]. "Ursus americanus". IUCN Red List of Threatened Species. 2016: e.T41687A45034604. doi:10.2305/IUCN.UK.2016-3.RLTS.T41687A45034604.en.
Hamilton, S. G.; Derocher, A. E. (2019). "Assessment of global polar bear abundance and vulnerability". Animal Conservation. 22 (1): 83–95. doi:10.1111/acv.12439.
"Fossilworks: Ursidae". Paleobiology Database. University of Wisconsin–Madison. Retrieved December 17, 2021.
García, N.; Santos, E.; Arsuaga, J. L.; Carretero, J. M. (December 2007). "Endocranial morphology of the Ursus deningeri von Reichenau 1904 from the Sima de los huesos (Sierra de Atapuerca) middle Pleistocene site". Journal of Vertebrate Paleontology. 27 (4): 1007–1017. doi:10.1671/0272-4634(2007)27[1007:EMOTUD]2.0.CO;2.
Garcı́a, N.; Arsuaga, J. L. (June 15, 2001). "Ursus dolinensis4: a new species of Early Pleistocene ursid from Trinchera Dolina, Atapuerca (Spain)". Comptes Rendus de l'Académie des Sciences, Série IIA. 332 (11): 717–725. doi:10.1016/S1251-8050(01)01588-9.
Pérez-Hidalgo, T.; José, T. (1992). "The European descendants of Ursus etruscus C. Cuvier (Mammalia, Carnivora, Ursidae)". Boletín del Instituto Geológico y Minero de España. 103 (4): 632–642.
Rabeder, G.; Hofreiter, M.; Nagel, D.; Withalm, G. (January 2004). "New taxa of alpine cave bears (Ursidae, Carnivora)". Cahiers scientifiques-Muséum d'histoire naturelle de Lyon. 2 (2): 49–67.
Kurtén, B. (1964). "The evolution of the polar bear, Ursus maritimus Phipps". Acta Zoologica Fennica. 108: 1–26. hdl:10138/37762.
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