Juniperus macrocarpa (large-fruited juniper, syn. J. oxycedrus subsp. macrocarpa (Sibth. & Sm.) Ball) is a species of juniper, native across the northern Mediterranean region from southwestern Spain[1] east to western Turkey and Cyprus, growing on coastal sand dunes from sea level up to 75 metres (246 feet) in altitude.[2][3]
Foliage and immature cones, Paros Island, Greece
Species of conifer
Juniperus macrocarpa
Juniperus macrocarpa in sand dune habitat, Paros Island, Greece
It is a spreading shrub2–5m (6+1⁄2–16+1⁄2ft) tall, rarely a small tree up to 14m (46ft) tall. The leaves are broad lanceolate, produced in whorls of three, green, 12–20 millimetres (1⁄2–3⁄4in) long and 2–3mm broad, with a double white stomatal band split by a green midrib on the inner surface. It is dioecious, with separate male and female plants. The seed cones are berry-like, green ripening in 18 months to orange-red with a variable pink waxy coating; they are spherical, 12–18mm diameter, and have six fused scales in two whorls, three of the scales with a single seed. The seeds are dispersed when birds eat the cones, digesting the fleshy scales and passing the hard seeds in their droppings. The pollen cones are yellow, 2–3mm long, and fall soon after shedding their pollen in late winter.[2][3][4]
Despite its distinct morphology with large cones and broad leaves more like those of Juniperus drupacea, it has often been treated as a subspecies of Juniperus oxycedrus,[3] though recent genetic studies[2][5][6] have shown its DNA is distinct from that of J. oxycedrus.
References
Muñoz-Reinoso, José Carlos (6 May 2004). "Diversity of maritime juniper woodlands". Forest Ecology and Management. 192 (2–3): 267–276. doi:10.1016/j.foreco.2004.01.039.
Adams, R. P. (2004). Junipers of the World. Trafford. ISBN1-4120-4250-X
Farjon, A. (2005). Monograph of Cupressaceae and Sciadopitys. Royal Botanic Gardens, Kew. ISBN1-84246-068-4
Adams, R. P. (2000). Systematics of Juniperus section Juniperus based on leaf essential oils and RAPD DNA fingerprinting. Biochem. Syst. Ecol. 28: 515-528. available online (pdf file)
Adams, R. P., Morris, J. A., Pandey, R. N., & Schwarzbach, A. E. (2005). Cryptic speciation between Juniperus deltoides and J. oxycedrus (Cupressaceae) in the Mediterranean. Biochem. Syst. Ecol. 33: 771-787. available online (pdf file)
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