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Cordia subcordata is a species of flowering tree in the borage family, Boraginaceae, that occurs in eastern Africa, South Asia, Southeast Asia, northern Australia and the Pacific Islands (including Hawaii).[2] The plant is known by a variety of names including beach cordia, sea trumpet, and kerosene wood, among others.

Cordia subcordata
Flower, fruit
Conservation status

Least Concern  (IUCN 3.1)[1]
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Boraginales
Family: Boraginaceae
Genus: Cordia
Species:
C. subcordata
Binomial name
Cordia subcordata

Names


Other names for the species include kanawa, tou, kou, mareer, manjak, snottygobbles, glueberry, narrow-leafed bird lime tree, In Java and Madura, it is known as kalimasada, purnamasada, or pramasada; Javanese folklore consider the tree to contain spiritual power.[3] In the Marshall Islands it is known as kono.[4]


Distribution


This species has a very wide range from the east coast of Africa west throughout tropical Asia and Oceania, as far west as Hawaii. This distribution was achieved due to special characteristics of its fruit allowing for successful oceanic dispersal.[5] Prior to 2001, C. subcordata was considered to be a Polynesian introduction to Hawaii by many authorities, but subfossil evidence from Makauwahi Cave in Kauai indicates that it was an abundant species in Hawaiian lowland forests well before humans arrived, confirming its status as an indigenous species.[6]


Description


C. subcordata grows to 7–10 m (23–33 ft) at maturity, but may be as tall as 15 m (49 ft). It has ovate leaves that are 8–20 cm (3.1–7.9 in) and 5–13 cm (2.0–5.1 in) wide.[7]


Flowers


The tubular flowers of C. subcordata are 2.5–4 cm (0.98–1.57 in) in diameter and form cymes or panicles.[7] Petals are orange and the sepals are pale green. Blooming occurs throughout the year, but most flowers are produced in the spring.[8]


Fruit


C. subcordata produces brown fruit year round. They are spherical, 2–3 cm (0.79–1.18 in) long, and woody when mature. Each fruit contains four or fewer seeds that are 10–13 mm (0.39–0.51 in) long. The fruit are buoyant and may be carried long distances by ocean currents.[7]


Habitat


C. subcordata is a tree of the coasts, found at elevations from sea level to 30 m (98 ft), but may grow at up to 150 m (490 ft). It grows in areas that receive 1,000–4,000 mm (39–157 in) of annual rainfall. C. subcordata prefers neutral to alkaline soils (pH of 6.1 to 7.4), such as those originating from basalt, limestone, clay, or sand. Allowable soil textures include sand, sandy loam, loam, sandy clay loam, sandy clay, clay loam, and clay.[7]


Uses


The seeds are edible and have been eaten during famine. C. subcordata burns readily, and this led to the nickname of "Kerosene Tree" in Papua New Guinea.[7] The wood of the tree has a specific gravity of 0.45, is soft, durable, easily worked,[8] and resistant to termites. In ancient Hawaiʻi kou wood was used to make ʻumeke (bowls), utensils, and ʻumeke lāʻau (large calabashes) because it did not impart a foul taste to food. ʻUmeke lāʻau were 8–16 litres (2–4 gal) and used to store and ferment poi. The flowers were used to make lei, while a dye for kapa cloth and aho (fishing lines) was derived from the leaves.[7]

In the western Solomon Islands, in Vanuatu, on Waya Island, and in Tonga, it is used for carving. On New Ireland, its wood is always used for the ceremonial entrances to men's houses.[9]


Threats


Despite its very wide distribution, this species faces threats in parts of its range. During the mid-late 19th century, herbivory by the kou moth (Ethmia nigroapicella) nearly extirpated this species on the Hawaiian Islands.[10] It is thought to be highly endangered in Sri Lanka, only persisting at a small number of sites that are under pressure from development.[5] Logging of mature trees to create carvings for the tourism industry is also thought to be a threat throughout much of Papuasia. In addition, it may potentially be threatened by storms and sea level rise.[1]


See also



References


  1. Oldfield, S. (2019). "Cordia subcordata". IUCN Red List of Threatened Species. 2019: e.T33199A67739216. doi:10.2305/IUCN.UK.2019-2.RLTS.T33199A67739216.en. Retrieved 19 November 2021.
  2. "Cordia subcordata". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA). Retrieved 2 March 2010.
  3. Bosbouwproefstation (1936). Korte mededeelingen - Cordia Subcordata. Cornell University. p. 107. Archived from the original on 31 July 2021. Retrieved 30 May 2015.
  4. "Native plants of the Marshalls". Republic of the Marshall Islands Biodiversity Clearing House Mechanism. RMI Office of Environmental Planning and Policy Coordination. 2 October 2008. Archived from the original on 19 June 2015. Retrieved 7 September 2019.
  5. Jayasuriya, A. H. Magdon; Gunatilleke, I. A. U. Nimal (18 September 2015). "A refugium for Cordia subcordata (Boraginaceae), a very rare and endangered plant in Sri Lanka and strategies for its conservation". Ceylon Journal of Science (Biological Sciences). 44 (1): 67. doi:10.4038/cjsbs.v44i1.7343. ISSN 0069-2379.
  6. Burney, David A.; James, Helen F.; Burney, Lida Pigott; Olson, Storrs L.; Kikuchi, William; Wagner, Warren L.; Burney, Mara; McCloskey, Deirdre; Kikuchi, Delores; Grady, Frederick V.; Gage, Reginald (November 2001). "FOSSIL EVIDENCE FOR A DIVERSE BIOTA FROM KAUA'I AND ITS TRANSFORMATION SINCE HUMAN ARRIVAL". Ecological Monographs. 71 (4): 615–641. doi:10.1890/0012-9615(2001)071[0615:FEFADB]2.0.CO;2. ISSN 0012-9615.
  7. Friday, J. B.; Dana Okano (April 2006). "Cordia subcordata (kou)" (PDF). Species Profiles for Pacific Island Agroforestry. The Traditional Tree Initiative. Archived (PDF) from the original on 16 January 2009. Retrieved 21 February 2009.
  8. Allen, James A. (1 January 2003). "Cordia subcordata Lam". Tropical Tree Seed Manual. Reforestation, Nurseries & Genetics Resources. Archived from the original (PDF) on 16 January 2009. Retrieved 24 February 2009.
  9. Pawley, Andrew; Osmond, Meredith (eds). 2008. The lexicon of Proto Oceanic: The culture and environment of ancestral Oceanic society Archived 31 July 2021 at the Wayback Machine. Volume 3: Plants Archived 4 February 2019 at the Wayback Machine. Pacific Linguistics 599. Canberra: Pacific Linguistics, Australian National University.
  10. Swezey, OH (1944). "The Kou Moth, Ethmia colonella Walsm., in Hawaii" (PDF). Hawaiian Entomological Society.


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