Cleistocactus is a genus of flowering plants in the cactus family Cactaceae, native to mountainous areas - to 3,000 m (9,843 ft)[1] - of South America (Peru, Uruguay, Bolivia and Argentina). The name comes from the Greek kleistos meaning closed because the flowers hardly open.

This article needs additional citations for verification. (December 2019)

Cleistocactus
Borzicactus websteramus, syn. Cleistocactus webstramus
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Order:	Caryophyllales
Family:	Cactaceae
Subfamily:	Cactoideae
Genus:	Cleistocactus Lem.
Species
See text

The stems of these cacti are tall, mostly slender and often many-branched with numerous ribs with closely set areoles and spines. The flowers are tubular and the tips hardly open with only the style and stamens usually protruding.

Species

Image	Scientific name	Distribution
	Cleistocactus acanthurus (Vaupel) D.R.Hunt	Peru
	Cleistocactus baumannii (Lem.) Lem.	Argentina, Paraguay, Bolivia, Uruguay
	Cleistocactus brookeae Cárdenas	Bolivia
	Cleistocactus buchtienii Backeb.	Bolivia
	Cleistocactus candelilla Cárdenas	Bolivia
	Cleistocactus chotaensis F.A.C.Weber ex Rol.-Goss.	Peru
	Cleistocactus clavispinus (Rauh & Backeb.) Ostolaza	Peru
	Cleistocactus dependens Cárdenas	Bolivia
	Cleistocactus ferrarii R.Kiesling	Argentina
	Cleistocactus fieldianus (Britton & Rose) D.R.Hunt	Peru (Amazonas, Ancash, Huanuco, Junin, Lima, Pasco, San Martin, Ucayali)
	Cleistocactus granditessellatus (Rauh & Backeb.) Leuenb.
	Cleistocactus grossei Backeb.	Paraguay
	Cleistocactus hildegardiae F.Ritter	Bolivia
	Cleistocactus hyalacanthus (K.Schum.) Rol.-Goss.	Argentina, Bolivia
	Cleistocactus hystrix (Rauh & Backeb.) Ostolaza	Peru
	Cleistocactus icosagonus (Kunth) F.A.C.Weber ex Rol.-Goss.	Ecuador, Peru
	Cleistocactus laniceps (K.Schum.) Rol.-Goss.	Bolivia
	Cleistocactus leonensis Madsen	Ecuador
	Cleistocactus longiserpens Leuenb.	Peru
	Cleistocactus luribayensis Cárdenas	Bolivia
	Cleistocactus micropetalus F.Ritter	Bolivia
	Cleistocactus morawetzianus Backeb.	Peru
	Cleistocactus muyurinensis F.Ritter	Bolivia
	Cleistocactus orthogonus Cárdenas	Bolivia
	Cleistocactus pachycladus (Rauh & Backeb.) Ostolaza	Peru
	Cleistocactus palhuayensis F.Ritter & Shahori	Bolivia
	Cleistocactus paraguarensis F.Ritter	Paraguay
	Cleistocactus parapetiensis Cárdenas	Bolivia
	Cleistocactus parviflorus (K.Schum.) Rol.-Goss.	Bolivia
	Cleistocactus peculiaris (Rauh & Backeb.) Ostolaza	Peru
	Cleistocactus piraymirensis Cárdenas	Bolivia
	Cleistocactus plagiostoma (Vaupel) D.R.Hunt	Peru
	Cleistocactus pungens F.Ritter	Peru
	Cleistocactus reae Cárdenas	Bolivia
	Cleistocactus ritteri Backeb.	Bolivia
	Cleistocactus roezlii (Haage ex K.Schum.) Backeb.	Peru
	Cleistocactus roseiflorus (Buining) G.D.Rowley
	Cleistocactus samaipatanus (Cárdenas) D.R.Hunt	Bolivia
	Cleistocactus sepium (Kunth) F.A.C.Weber ex Rol.-Goss.	Ecuador
	Cleistocactus serpens (Kunth) F.A.C.Weber ex Rol.-Goss.	Ecuador, Peru
	Cleistocactus sextonianus (Backeb.) D.R.Hunt	Peru
	Cleistocactus smaragdiflorus (F.A.C.Weber) Britton & Rose	Argentina
	Cleistocactus strausii (Heese) Backeb.	Bolivia, Argentina
	Cleistocactus sulcifer (Rauh & Backeb.) Leuenb.	Peru
	Cleistocactus tarijensis Cárdenas	Bolivia
	Cleistocactus tenuiserpens Rauh & Backeb.	Peru
	Cleistocactus tominensis (Weing.) Backeb.	Bolivia
	Cleistocactus tupizensis (Vaupel) Backeb.	Bolivia
	Cleistocactus variispinus F.Ritter	Bolivia
	Cleistocactus vulpis-cauda F.Ritter & Cullmann	Bolivia
	Cleistocactus winteri D.R.Hunt	Bolivia
	Cleistocactus xylorhizus (F.Ritter) Ostolaza	Peru

Natural Hybrids

• Cleistocactus × crassiserpens Rauh & Backeb.(Cleistocactus icosagonus × Cleistocactus serpens).

Synonyms

The following genera have been brought into synonymy with this genus:

• Akersia Buining
• Bolivicereus Cárdenas
• Borzicactella H.Johnson ex F.Ritter
• Borzicactus Riccob
• Borzicereus Fric & Kreuz. (orth. var.)
• Cephalocleistocactus F.Ritter
• Cleistocereus Fric & Kreuz. (orth. var.)
• Clistanthocereus Backeb.
• Demnosa Fric
• Gymnanthocereus Backeb.
• Hildewintera F.Ritter
• Loxanthocereus Backeb.
• Maritimocereus Akers
• Pseudoechinocereus Buining (nom. inval.)
• Seticereus Backeb.
• Seticleistocactus Backeb.
• Winteria F.Ritter
• Winterocereus Backeb.

• 
  Cleistocactus strausii
• 
  Cleistocactus micropetalus F. Ritter, photo taken at Bedugul Botanical Garden, Bali, Indonesia

References

• Innes C, Wall B (1995). Cacti, Succulents and Bromaliads. Cassell & The Royal Horticultural Society.
• The species list is referenced from http://www.cactiguide.com/ which is in turn referenced from several books which are listed on that site. The principal book listed here is The Cactus Family by Edward F. Anderson.

External links

Wikimedia Commons has media related to Cleistocactus.

• (in French) photos on www.AIAPS.org
• (in English) photos on www.cactiguide.com

На других языках

[es] Cleistocactus

[ru] Клейстокактус