Quercus pubescens, the downy oak or pubescent oak, is a species of white oak (genus Quercus sect. Quercus) native to southern Europe and southwest Asia, from northern Spain (Pyrenees) east to the Crimea and the Caucasus. It is also found in France and parts of central Europe.

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Downy oak
A mature tree
Leaves
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Rosids
Order:	Fagales
Family:	Fagaceae
Genus:	Quercus
Subgenus:	Quercus subg. Quercus
Section:	Quercus sect. Quercus
Species:	Q. pubescens
Binomial name
Quercus pubescens Willd.
Distribution map
Synonyms
List Eriodrys lanata Raf. Quercus adjecta Gand. Quercus admixta Gand. Quercus aegilops Mill. Quercus amplifolia Guss. Quercus appenina Lam. Quercus aspera Bosc Quercus asperata Pers. Quercus bacunensis Vuk. Quercus banja Endl. Quercus bellojocensis Gand. Quercus brachyloba Jord. Quercus brachyphylla Kotschy Quercus brachyphylloides Vuk. Quercus brandisii (Vuk.) Vuk. Quercus brevifolia Kotschy ex A.DC. Quercus buccarana Vuk. Quercus budayana Haberle ex Heuff. Quercus budensis (Borbás) Borbás Quercus collina Schleich. ex Endl. Quercus conglomerata Pers. Quercus croatica Vuk. Quercus cupaniana] Guss. Quercus dalmatica Radic Quercus diversifrons Borbás Quercus erythrolepis (Vuk.) Vuk. Quercus humilis Mill. Quercus ilicifolia Koord. & Valeton ex Seemen Quercus lacinifolia Vuk. Quercus laciniosa Boreau Quercus lanuginosa (Lam.) Thuill. Quercus macrostipulata Guss. ex Parl. Quercus menesiensis Kit. Quercus microbalanos Boreau Quercus microlepis Vuk. Quercus oxycarpa Raddi Quercus pinnatifida C.C.Gmel. Quercus pseudoaegilopsis Petz. & G.Kirchn. Quercus rufa Vuk. Quercus schulzei (Vuk.) Vuk. Quercus sectifolia Vuk. Quercus stenolepis Vuk. Quercus subspicata (A.Camus) C.Vicioso Quercus sulcata Vuk. Quercus susedana Vuk. Quercus tenorei (A.DC.) Borzí Quercus torulosa (Vuk.) Raddi Quercus virgiliana (Ten.) Ten. Quercus vukotinocicii Borbás Quercus anatolica O.Schwarz Quercus crispata Steven Quercus subpyrenaica Villar

Description

Quercus pubescens is a medium-sized deciduous tree growing up to 20 metres (66 feet). Forest-grown trees grow tall, while open-growing trees develop a very broad and irregular crown. They are long-lived, to several hundred years, and eventually grow into very stout trees with trunks up to 2 m (6+1⁄2 ft) in diameter. Open-grown trees frequently develop several trunks. The bark is very rough, light gray and divided into small flakes. Large trees develop very thick whitish bark cracked into deep furrows, similar to the pedunculate oak but lighter in colour.

The twigs are light purple or whitish, with tomentum. The buds are small (3–6 millimetres or 1⁄8–1⁄4 inch) and blunt, light brown. The leaves are leathery usually 4–10 centimetres (1+1⁄2–4 in) long (rarely to 13 cm) and 3–6 cm wide, usually widest beyond the middle. The leaves group at the ends of twigs. The upper leaf surface is dark green and rough, the lower light green. Both leaf surfaces are covered with minute pubescence which is sometimes lost in older leaves by late summer. The young expanding leaves are whitish or pinkish with very soft tomentum. The leaf shape is very variable, divided into 3–7 pairs of deep or shallow lobes, which are usually divided into a few sublobes. The lobes are usually blunt, rarely sharp. The apex is usually wide and round. The base of the leaf is heart shaped, widely rounded or sometimes pointed. The petioles are 4–15 mm (rarely to 22 mm) long, stout and pubescent. The leaves are persistent late into the autumn, remaining green up to early winter. They eventually turn russet or brown and fall off.

The Quercus pubescens acorns are light brown to yellow, 8–20 mm long, usually thin and pointed. The acorn cups are light gray to almost white, with pointed, overlapping scales, covered with tomentum. The acorn stalks are thick and pubescent, up to 2 cm long. The acorns usually occur in groups of 2–5 on the same stalk.

Subspecies

Three subspecies are accepted by Flora Europaea:

• Quercus pubescens subsp. pubescens – central and southern Europe.
• Quercus pubescens subsp. anatolica O.Schwarz – southwest Asia, southeast Europe.
• Quercus pubescens subsp. palensis (Palassou) O.Schwarz – northern Spain, Pyrenees.

Habitat

Downy oaks typically grow in dry, lime-rich soils. It is a submediterranean species, growing from the coastline to deep in the continent. Its optimum is in the submediterranean region, characterized by hot dry summers and cool winters with little rainfall. In western and central Europe, downy oak is confined to areas with submediteranean microclimate (gorges, sandplains, steppe slopes) or to coastlines of former lakes.

References

• Flora Europaea: Quercus pubescens
• Bean, W. J. (1976). Trees and shrubs hardy in the British Isles 8th ed., revised. John Murray.
• Rushforth, K. (1999). Trees of Britain and Europe. HarperCollins ISBN 0-00-220013-9.
• (in French) Chênes: Quercus pubescens
• Quercus pubescens and Quercus virgiliana - information, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)

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На других языках

[es] Quercus humilis

[ru] Дуб пушистый