Currently there are about 40 extant families of Lacertilia. These vary considerably, e.g. in shades, colours, and sizes. For example, the largest representative among Geckos, the New Caledonian giant gecko (Rhacodactylus leachianus), has a length of up to 36cm (14in), while the largest species in the family Varanidae, Komodo dragon (Varanus komodoensis), has a length up to 3 metres (10ft), and a body mass of 70kg (154lbs).
Largest extant lizards
Here are the 15 largest extant lizards based on the most reliable measurements for size, including length and weight. These include family types Varanidae, Iguanidae and Teiidae exceeding 9kg (20lbs) in mass.
Makassar salfin lizard is usually regarded as the largest representative of family Agamidae, reaching a length of 1.2m (3.9ft).
The largest representatives in this group are species in the genus Hydrosaurus, which often exceed 1 metre (3.3ft) in length.[24]
The largest species – Makassar salfin lizard (Hydrosaurus microlophus) and Sulawesi black salfin lizard (Hydrosaurus celebensis) growing a length of 1.2(3.9ft) and 1.13m (3.7ft) respectively.[24]
A Weber's salfin lizard (Hydrosaurus weberi), Philippine salfin lizard (Hydrosaurus pustulatus) and Amboina salfin lizard (Hydrosaurus amboinensis) (previously counts the largest species in this group), have a length of 1m (3.3ft).[24]
The Australian water dragon (Intellagama lesueurii) and Chinese water dragon (Physignathus cocincinus) can also exceed this length.[25][26][27]
The frilled lizard (Chlamydosaurus kingii) has length up to 85cm (2.79ft) and mass up to 870g (1.92lb),[28] probably even 900g (2lb)[29]
Uromastyx aegyptia is one of heaviest representatives among agamas, with length up to 76cm (30in)[30] and mass 2kg (4.4lbs).[31]
A central bearded dragon (Pogona vitticeps) and eastern bearded dragon (Pogona barbata) have a length of 60cm (24in),[32][33][34] and are the largest in their genus.[34]
American legless lizards (Anniellidae)
See also: Anniellidae
The largest american legless lizards is the Californian legless lizards (Anniella pulchra) attaining 18cm (7.1in) long.
Glass lizards (Anguidae)
See also: Anguidae
Sheltopusik from Southern Europe to Central Asia, is the largest legless lizard, reaching a length of 1.2 m (3.9 ft)
Many representatives of this family are legless lizards. Among these the largest species, and also the largest legless lizard in the world, is the sheltopusik (Pseudopus apodus), reaching a maximum length of more 135cm (4.43ft) and a mass in 1.1kg (2.4lb).[35]
The eastern glass lizard (Ophisaurus ventralis) and slender glass lizard (Ophisaurus attenuatus) may reach lengths of up to 108cm (3.54ft)[36] and 107cm (3.51ft)[37] respectively.
However, this family also has lizards with legs, among which the largest is the Texas alligator lizard (Gerrhonotus infernalis) growing to from 45.7cm (18in) till 64cm (25.2in)[38][39] in different sources.
Chameleons (Chamaeleonidae)
See also: Chamaeleonidae
Of all chameleons, the largest is the Parson's chameleon (Calumma parsonii), reaching a length of 68cm (27in).[40][41] But the longest species is the Malagasy giant chameleon (Furcifer oustaleti). It has a length of up to 68.5cm (27in).[42]
Other large chameleons are the veiled chameleon (Chameleo calyptratus) and Meller's chameleon (Trioceros melleri), with lengths of up to 61cm (24in)[43][44] Exceptionally large specimens of the latter have reputedly reached over 76cm (30in) and mass 600 g (1.3lb), although this is unlikely.[45]
Spinytail lizards (Cordylidae)
See also: Cordylidae
Many species in this family are small to medium-sized lizards that range in length from 6 to 30cm (2.4 to 11.8inches).[46] There are large species in the genus Smaug, the largest of which is a giant girdled lizard (Smaug giganteus), with snout-vent length up to 20.5cm (8.07in)[47] and total length up to roughly 40cm (15.74in).[48]
Casquehead lizards (Corytophanidae)
See also: Corytophanidae
Basilisks are quite large lizards, especially the plumed basilisk
Casquehead lizards are quite large lizards. In particular, the genus Basiliscus typically measures 70–75cm (28–30in).[49][50][51] But the largest species in this group is the plumed basilisk (Basiliscus plumifrons), frequently growing to 91cm (36in).[52][53]
Another large species in this group is the eastern casquehead iguana (Laemanctus longipes) reaching up to 70cm (27.6in) including its long, thin tail.
Collared lizards (Crotaphytidae)
See also: Crotaphytidae
In this family it is not known exactly which species is the largest, but according to current data the largest is the reticulated collared lizard (Crotaphytus reticulatus) with length up to 41cm (16in)[citation needed] and mass 78.77g (2.8oz).[54]
Anoles (Dactyloidae)
See also: Dactyloidae
The knight anole is the largest anole, over half a meter in length
Large species in this group are usually crown giant anolis ecomorphs[55][56][57] The species generally considered the largest is the knight anole (Anolis equestris), which can reach up to 51cm (20in) in total length, 19cm (7.5in) in snout-vent length and weigh 137g (4.83oz).[58][59]
However, in the equester species complex there are also species reaching the same length and even larger. These include A. baracoae, A. luteogularis, A. noblei and A. smallwoodi.[60][61][62][63]
Geckos (Gekkota)
See also: Gekkota
The New Caledonian giant gecko is the largest gecko, weighing up to almost 300g (10.5oz)
Species of geckos belong to different groups. For example, many species of geckos are in family Gekkonidae, but there are very large species in the family Diplodactylidae. For example, to them belongs the largest gecko species New Caledonian giant gecko (Rhacodactylus leachianus), which can be 36–43.18cm (14–17in) long and mass 212–279g (7.5–9.8oz)[64][65]
Other representatives of the same genus as the aforementioned gecko can also reach huge sizes. These include gargoyle gecko (Rhacodactylus auriculatus), greater rough-snouted giant gecko (Rhacodactylus trachyrhynchus) and lesser rough-snouted giant gecko (Rhacodactylus trachycephalus).[66][67][68] As well as former species in this genus: crested gecko (Correlophus ciliatus), Sarasins' giant gecko (Correlophus sarasinorum) and Bavay's giant gecko (Mniarogekko chahoua).[69][70][71]
A extant member of the genus Hoplodactylus, Duvaucel's gecko (Hoplodactylus duvaucelii), in which be present a largest gecko ever lived (see the largest extinct lizards) can also reach large sizes. It can reach total length 30cm (12in)[72] with a snout-vent length (SVL) up to 16cm (6.3in), and weigh up to 120 grams (4.2oz).[73]
There are also large species in the family Gekkonidae. These include tokay gecko (Gekko gekko), Smith's green-eyed gecko (Gekko smithii) and Andaman giant gecko (Gekko verreauxi), which can reach and even exceed 30cm (12in).[74][75][76][77][78]
Another genus with large geckos is Uroplatus. The largest representative in this genus is the giant leaf-tail gecko (Uroplatus giganteus), which can reach a snout–vent length of 20cm and a total length of 32.2cm.[79] Another species that grows almost as large is the common flat-tail gecko (Uroplatus fimbriatus), which can exceed 30cm (12in) and more.[80][81]
The genus Phelsuma in subfamily Uroplatinae also has large species. Standing's day gecko (Phelsuma standingi) is one of the largest day geckos, reaching a length of about 25.4cm (10.0in)[82] and Round Island day gecko (Phelsuma guentheri), Madagascar giant day gecko (Phelsuma grandis) and Phelsuma madagascariensis can reach even more impressive lengths of 30cm (12in) or more.[83][84][85]
Outside of the families Diplodactylidae and Gekkonidae, there are large geckoes among the Eublepharidae. The African fat-tailed gecko (Hemitheconyx caudicinctus) reaches a length of up to 23cm (9.1in),[86] and a mass of 75g (2.6oz).[87] The leopard gecko (Euplepharis macularis) reaches even large sizes – up to 28cm (11in) long with a mass of 60–80g (2.1–2.84oz).[88]
The largest species of a family Phyllodactylidae, the giant wall gecko (Tarentola gigas) growing 30cm (12in) in length,[89] and 15.5cm (6.1in) in SVL.[90]
Plated lizards (Gerrhosauridae)
See also: Gerrhosauridae
The largest species in this family is the giant plated lizard (Matobosaurus validus), with a total length of up to 69cm (27in)[91][92] or even 75cm (29.5in),[93] with a snout-vent length (SVL) of 28cm (11in).[91] Another large plated lizard is the Sudan plated lizard (Broadleysaurus major), with a length of about 51–60cm (20–24in).[94]
Beaded lizards and Gila monster (Helodermatidae)
See also: Gila monster and Helodermatidae
The Mexican beaded lizard is the largest species in family Helodermatidae, reaching a length of up to 91cm (36in)
This family includes large and stocky lizards, including the infamous Gila monster (Heloderma suspectum), which reaches a total length up to 60cm (23.6in) with a snout-vent length (SVL) of 36cm (14.1in)[7] and a body mass from 1kg (2.2lb)[7] to 2.2kg (4.9lb)[citation needed] depending on the source. But despite the stocky and massive body of this lizard, it is not the largest representative of the family. Greatly exceeding it in size are the Mexican beaded lizard (Heloderma horridum) and Rio Fuerte beaded lizard (Heloderma exasperatum), with a total length of up to 91cm (36in),[7][95] a SVL of up to 47cm (18.5in)[7] and a mass from 2kg (4.4lb)[7] to 4kg (8.8lb)[96][97][95] depending on the source.
Iguanas (Iguanidae)
See also: Iguanidae
The blue iguana is the heaviest iguanid and one of the largest extant lizards
Iguanas are very large lizards, some of which can reach lengths of up to 2m (6.6ft) and mass more than 10kg (22lb). They are the largest lizards after some large species of monitor lizards, and the largest lizards in the New World. Iguanas vary considerably in size and form, but even the smallest lizards in this family are still quite large. For example, the desert iguana (Dipsosaurus dorsalis) can reach a length of up to 61cm (24in).[98] The largest species of the genus Sauromalus is the San Esteban chuckwalla (Sauromalus varius), up to 76cm (27in) long and 1.4kg (3.1lb) in weight,[99] while the second-largest is the Angel Island chuckwalla (Sauromalus hispidus), up to 64cm (25.2in) long.[99]
Many sources describe the green iguana (Iguana iguana) as the largest iguanid, often reaching lengths up to 1.5 metres (4.91ft) and masses of 4kg (8.8lb),[100] and with a maximum length of 2m (6.6ft) and a mass of 8kg (18lb)[20][101] and in some cases even 9.1kg (20lb).[20] However, the heaviest species in this family is the blue iguana (Cyclura lewisi), with a total length of up to 1.5m (4.91ft), a SVL of 51–76cm (30in) and a mass of up to 14kg (31lb)[10][11] It is the eighth-heaviest and largest extant lizard.[12]
The Galapagos land iguana is a very massive lizard weighing more than 10kg (22lb) and is one of the largest extant lizards
Other large species in this family include the Galapagos land iguana (Conolophus subcristataus), with a length of about 1.5m (4.91ft) and a mass of up to 13kg (29lb).[13][14] It is the second-heaviest iguanid after the blue iguana and the ninth-heaviest and largest lizard in the world.[12] Another large species from the same genus is the Santa Fe land iguana (Conolophus pallidus), reaching a SVL of 56.4cm (22.2in) and a mass of 7.2kg (16lb).[102] The Galapagos pink land iguana (Conolophus marthae) have snout-vent length 57.5cm (22.6in) and the mass of 8kg (18lb).[102]
The marine iguana (Amblyrhynchus cristatus) is the tenth-largest extant lizard in the world,[12] and the largest reptile on Galapagos Islands after the Galapagos land iguana, not including turtles reaching a maximum total length of 1.4m (4.59ft), a SVL of from 12 till 56cm (from 4.72 till 22in)[16][17] and a mass of from 1 to 12kg (2.2 to 26.5lb)[15] depending of islands.
Aforementioned genus Cyclura includes other very large iguanas, such as Anegada rock iguana (Cyclura pinguis) reaching a snout-vent length (SVL) of 56cm (22in)[103] and a mass 7.75kg (17.1lb).[104] A second largest species in this genus and second longest iguanid overall, and also one of the largest lizards in the Caribbean, the Cuban iguana (Cyclura nubila) reaching a length in 1.6m (5.2ft) with a SVL in 74.5cm (29.3in).[104]
The rhinoceros iguana (Cyclura cornuta) has a length in 1.36m (4.5ft), with a SVL 56cm (22in) and a mass more 4.56–9kg (10.1–19.8lb).[104] Previously considered of the subspecies aforementioned rhinoceros iguana, the Mona ground iguana (Cyclura strejnegeri) exceeds a total length of 1.22m (4ft),[105] the SVL 53.5cm (21.1in), mass of 6.4kg (14lb)[104] and is the largest native terrestrial lizard in Puerto-Rico.
At least two subspecies (Cyclura cychlura cychlura and Cyclura cychlura inornata) of the Northern Bahamian rock iguana (Cyclura cychlura) can be over 1.5m (4.9ft) in length.[106][107][108][109]
A member of the genus Iguana, lesser Antillean iguana (Iguana delicatissima) can reach SVL 43.4cm (17.1in), and mass 3.5kg (7.7lb).[110]
The black iguana (Ctenosaura similis) and western spiny-tailed iguana (Ctenosaura pectinata) are the largest species in the genus Ctenosaura, reaching a length in 1.3m (4.3ft).[111][112] The longest is Mexican spiny-tailed iguana (Ctenosaura acanthura) with a length of 1.4m (4.6ft).[113]
True lizards (Lacertidae)
See also: Lacertidae
Gran Canaria giant lizard is the largest member of a family Lacertidae, exceeding a length of 80cm (31in)
This family includes 300 diverse species. Among the largest is Gran Canaria giant lizard (Gallotia stehlini), reaching a length of 80cm (31in),[114][115] with a SVL of 26.5cm (10.4in)[116] and a mass of almost 1.7kg (3.7lb).[117] A subspecies the same genus – El Hierro giant lizard (Gallotia simonyi machadoi), perhaps also is one of the largest lacertid, and able reach a length of 60cm (24in).[citation needed] A Tenerife lizard (Gallotia galloti) have a total length in 44cm (17in)[citation needed] and a SVL of 14.5cm (5.7in).[118] Another large lizard in this genus are La Gomera giant lizard (Gallotia brovoana), reaching a length of almost 50cm (20in) and a SVL in 19cm (7.5in)[119]
Estimated to reaching the snout-vent length of 40–44cm (16–17in) and a mass in 5kg (11lb),[120] La Palma giant lizard (Gallotia auaritae) perhaps the largest living lacertid outsized even Gran Canaria giant lizard (Gallotia stehlini). However, on the other hand, it is very little data to confirm this, and therefore the aforementioned Gran Canaria giant lizard is usually considered the largest representative of its family.
The longest member of this family is ocellated lizard (Timon lepidus), having a length of 90cm (35in),[121] with a SVL 26cm (10in)[122] and a mass 0.5kg (18oz).[123] A representative the same genus, the Moroccan eyed lizard (Timon tangitanus) can have a length of 70cm (28in),[citation needed] and is one of the longest members of the this family.
The European green lizard (Lacerta viridis), Iberian emerald lizard (Lacerta schreiberi) and western green lizard (Lacerta bilineata) can exceed a length 40cm (16in)[124][125] and more, and with a SVL at least 13cm (5.1in).[126] The largest species in this group is Balkan green lizard (Lacerta trilineata), reaching a length of 50cm (20in), and a SVL of 16cm (6.3in).[127]
Earless monitor lizards (Lanthanotidae)
See also: Lanthanotidae
Only living species this family – earless monitor lizard (Lanthanotus borneensis) typically have a snout-vent length (SVL) of about 20cm (8in),[128] and a total length of about 40cm (16in).[129] The longest recorded in the wild had length in 44.1cm (17.4in).[130] A specimen collected in the 1960s had a total length of 51cm (20in),[131] and near the time of its death an individual kept at the Bronx Zoo from 1968 to 1976 had a total length of 47cm (18.5in) and weighed 209g (7.4oz), but it was highly obese.[132]
Curly-tailed lizards (Leiocephalidae)
See also: Leiocephalidae
The largest curly tailed lizards Leiocephalus carinatus Adults may attain a snout to vent length (SVL) of 10.5cm (4.1in), or a total length, including the tail, of 26cm (10in). The dorsal scales are keeled and pointed. L. carinatus resembles lizards of the genus Sceloporus, but with the tail usually curled upward, especially when the lizard is in a horizontal position on rocks or on the ground.[133]
Leiosaurids (Leiosauridae)
See also: Leiosauridae
The Largest ever Enyalius catenatus The body mass can be up to 38.66g (1.364oz). The species is reproduces sexually and is gonochoric.[134]
Liolaemids (Liolaemidae)
See also: Liolaemidae
Members of the genus Liolaemus are vary considerably in size (45–100 millimetres or 1.8–3.9 inches snout–vent length) and weight (3–200 grams or 0.1–7.1 ounces). The largest species are L. chlorostictus (85mm max. SVL), L. dorbignyi (98.3mm max. SVL), L. duellmani (83mm max. SVL), L. forsteri (93mm max. SVL), L. foxi (82.9mm max. SVL), L. huayra (94.3mm max. SVL), L. inti (90.4mm max. SVL), L. jamesi (97.5mm max. SVL), L. melanogaster (91mm max. SVL), L. nigriceps (88.8mm max. SVL), L. orientalis (99.2mm max. SVL), L. patriciaiturrae (96.5mm max. SVL), L. puritamensis (90.8mm max. SVL), L. robertoi (84mm max. SVL), L. scrocchii (95mm max. SVL) and L. stolzmanni (92mm max. SVL).[135] The L. fabiani, L. filiorum, L. pachecoi, L. pleopholis, L. polystictus, L. robustus, L. thomasi, L. vallecurensis, L. vulcanus, and L. williamsi, L. austromendocinus, L. elongatus, L. gununakuna, L. petrophilus, and L. thermarum also may reach large sizes (around 10cm)[136][137]
Madagascan iguanas (Opluridae)
See also: Opluridae
The biggest species of Madagascan iguanas is Madagascar spotted spiny-tailed iguana (Oplurus quadrimaculatus), it can reach a length of 25–39 centimetres (9.8–15.4in).
Legless lizards (Pygopodidae)
See also: Pygopodidae
The largest legless lizard this family is the common scaly-foot (Pygopus lepidopodus) with a length of 70cm (28in),[138] a SVL of 24cm (9.4in)[139] with an weight of 225.95g (7.970oz)[140] and also the largest legless lizard in Australia[138] and biggest in the world overall, if don't counts some members the family Anguidae.
The another large member is Burton's legless lizard (Lialis burtonis) with a maximum length of 60cm (23.6in),[141] according to some information even 1.02m (3.3ft)[142][143] and a body length (snout-vent length) of 29cm (11in).[142][143][144]
Skinks (Scincidae)
See also: Scincidae
The Solomon Islands skink – largest skink reaching the length of 81cm (32in)
In this family very lot of species – about 1500. And many of species skinks have a snout-vent length (SVL) in 12cm (4.7in). Although, the largest member of this family – Solomon Islands skink (Corucia zebrata) can grows the length of 81cm (32in),[145][146] a SVL in 35cm (13.8in) and a mass of 0.8–0.9kg (1.8–2.0lb).[147][148] But the heaviest is the common blue-tongued skink (Tiliqua scincoides) reaching a length in 60cm (24in),[149][150][151] a SVL of 36cm (14.1in)[152][149] and the mass of 1kg (2.2lb).[153]
In the genus Tiliqua be present also other a large species of skinks. One of them is the subspecies Indonesian blue-tongued skink (Tiliqua gigas) – Merauke blue-tongued skink (Tiliqua gigas evanescens) growing a length of 66–76cm (26–30in),[151] thus not only being the longest species in the genus Tiliqua, but also the second by length in family Scincidae after Solomon Islands skink. The large sizes also may reach blotched blue-tongued lizard (Tiliqua nigrolutea) with a length of 35–50cm (14–20in),[154] according to some information even 59.6cm (23.5in),[151][150] a SVL of 32cm (13in) and a mass of 0.8kg (1.8lb),[155] Centralian blue-tongued skink (Tiliqua multifasciata) exceeding in length of 45cm (18in)[151] with a SVL of 28.9cm (11.4in),[156] western blue-tongued lizard (Tiliqua occipitalis) with a length of 50cm (20in)[157][150] and a SVL of 30.7cm (12.1in),[156] probably 32cm (13in),[158] shingleback lizard (Tiliqua rugosa) with a total length of 41cm (16in),[159] a SVL of 26–31cm (10–12in)[160] and a mass of 0.6 to 0.9kg (1.3 to 2.0lb)[161] and a Irian Jaya blue tongue skink (Tiliqua sp.) with total length of 38–76cm (15–30in).[162]
Previously considered of species in the genus Tiliqua is pink-tongued skink (Cyclodomorphus gerrardii) has a length of 45cm (18in)[163][164] and a SVL of 20cm (7.9in).[165]
The land mullet (Bellatorias major) is one of the largest skinks and can reach a SVL of 16–31cm (6.3–12.2in)[166] with a total length of 60cm (24in).[167] The representative the same genus – the major skink (Bellatorias frerei) also grows large sizes – a total length in 42cm (17in) with a SVL of 18cm (7.1in)[168]
The King's skink (Egernia kingii) is one of the longest skinks and can exceed a length of 55cm (22in)[153] with a SVL of 24.4cm (9.6in)[169] and weighing 220g (7.8oz).[170] Other large skinks are Cunningham's spiny-tailed skink (Egernia cunninghami) and yakka skink (Egernia rugosa), both of which reach a length in 40cm (16in).[171][172]
A representative of the genus Eumeces is Schneider's skink (Eumeces schneiderii) has a length of 36.5cm (14.4in) with a SVL in 16.5cm (6.5in).[173] Previously considered of species in the aforementioned genus – Kishinoue's giant skink (Plestiodon kishinouyei) is the largest species in its genus and may grows a length of 30–40cm (12–16in) with a SVL of 14.0–17.2cm (5.5–6.8in). Other large members in the genus Plestiodon are a broad-headed skink (Plestiodon laticeps) and a Great Plains skink (Plestiodon obsoletus) with a maximum length of 32.4cm (12.8in)[174] and 34.9cm (13.7in)[175] and a SVL 14.3cm (5.6in)[174] and 14cm (5.5in)[176] respectively.
The genus Chalcides includes many legless or almost legless skinks. The largest among these are the Italian three-toed skink (Chalcides chalcides) and the western three-toed skink (Chalcides striatus) which have a length of 48cm (19in) and 43cm (17in)[177] respectively. In the genus Acontias also are limbless skinks many of which are quite small lizards, but the largest among these is giant legless skink (Acontias plumbeus) at approximately reaching 40cm (16in) in length.[178]
The fire skink (Mochlus fernandi) is the largest member in its genus and also one of the largest African skinks, growing the length of 37cm (15in) and a SVL of 17cm (6.7in)[179]
Telfair's skink (Leiolopisma telfairii) the largest living skink in Mauritius, attains a size over 16cm (6.3in) in SVL[180] and may grows 30–40cm (12–16in) in length[181] with a maximum length of 45cm (18in) and a mass in 240g (8.5oz).[182] The its extinct relative is Mauritian giant skink (Leiolopisma mauritiana), which in twice excelled to length of its contemporary (see the largest extinct lizards) and was the biggest known skink of all time.
The terror skink (Phoboscincus bocourti) is a endangered skink and may reach 50cm (20in) in length[183] and 28cm (11in) in SVL[184][185] thus not only being one of the largest extant skinks, but the second biggest reptile on its island after monitor lizards, and so is an apex predator in its territory.[183]
Tegus and Teiids (Teiidae)
See also: Teiidae
Argentine black and white tegu as well as a red tegu, is the largest teiid and one of the heaviest lizards in the world reaching a mass of 7kg (15lb)
A many of teiids are quite small lizards. Although some members such as tegus are one of the biggest lizards in the world after monitor lizards and iguanas. The largest species this large family are Argentine black and white tegu (Salvator merianae) and red tegu (Salvator rufescens) attains more 1.2m (3.9ft) in a length and 6.8kg (15lb) in a mass. Argentine black and white tegu usually counts the largest species in its family with a length of 1.5m (4.91ft) and a mass of 7kg (15lb). Although, however, the red tegu, as more massive and bulky, can weigh more – the length is about 1.4m (4.6ft), and the weight is 9kg (20lb).[23]
Both species of genus Dracaena, the northern caiman lizard (Dracaena guianensis) and the Paraguay caiman lizard (Dracaena paraguayensis), can have a length of 1.2m (3.9ft) and a mass in 4.5kg (9.9lb).[186]
The largest member of the genus Tupinambis is the gold tegu (Tupinambis teguixin) with a maximum SVL in 40cm (16in).[187] Other large teiids this group are the cryptic golden tegu (Tupinambis cryptus) (maximum size: 391mm (15.4in) SVL with a 530mm (21in) tail), the Cusco tegu (Tupinambis cuzcoensis) (SVL of 247mm (9.7in) or larger) and the Maracaibo Basin tegu (Tupinambis zuliensis) (a 273mm (10.7in) SVL with a 537mm (21.1in) tail).[188]
Monitor lizards (Varanidae)
Body size in monitor lizards shows greater variation than in any other family of animals[189] with adult animals weighing between 23g (0.81oz) and 80,000g (176.3lb). The family includes both the largest living lizards and the largest lizards that have ever existed, yet about a third of the living species are dwarfs that seldom exceed 500g (18oz) in weight.[189] Such massive size disparity between species makes the group ideal model animals to study the effects of gigantism on ecology and physiology, but at present little of their potential in this field has been explored.[189] Although, species weighing around 1kg (2.2lb) are often found. This family includes the Komodo dragon, which is the largest of all extant lizards, with a length of 3 meters (10ft) and approximate weight of 113kg (249lb).[184] Some of the largest representatives of the Varanidae such as the Komodo dragon, crocodile monitor, perentie and lace monitor can count an apex predators. In the list of the largest lizards, monitors occupy the first seven places.[12]
Komodo dragon (Varanus komodoensis)
The Komodo dragon is the largest extant lizard with a maximum known mass of more 80kg (176.3lb), also perhaps the heaviest squamate
The Komodo dragon (Varanus komodoensis) is the largest living lizard in the world, with an average mass in 70kg (150lb) and 25kg (55lb) for males and females respectively.[190] The largest males from Komodo, may reach more 80kg (176.3lb). The largest specimen of Komodo dragon, a large male from Loh Liang National Park with length of 3.04m (10.0ft), a SVL of 1.54m (5.1ft) and a mass in 81.5kg (178.5lb).[2][3] As stated above, females are generally much smaller than males. The largest female weighed about 42kg (93lb).[191] The heaviest specimen in the wild weighed about 87.4kg (193lb).[2][3] The study noted that weights greater than 100kg (220lb) were possible but only after the animal had consumed a large food.[1][2][3] The largest accurately measured individual according to Gerard Wood for Guinness World Records was a specimen put on display at the Saint Louis Zoological Park, Missouri in 1937. This individual reportedly measured 3.13m (10.27ft) and weighed 165.5kg (365lb), although a large portion of this was likely undigested food. Once again captive animal are typically overweight to a certain degree so this weight is again much higher than what is achievable by wild individuals.[1] Thus, the Komodo dragon is not only the largest living lizard, but also the heaviest, if not the heaviest, then one of the heaviest representatives of the order Squamata (the only weight competitor is the green anaconda (Eunectes murinus) with a weight of about 97.5kg (215lb),[192] others are some species of pythons, but data are scarce) according to recorded data in the wild, and also second in mass after Burmese python (Python bivittatus) (mass – about 182.8kg[403lb])[193] in captive.
Asian water monitor (Varanus salvator)
Asian water monitor is the second-largest lizard with a length of more 3m (10ft) and a mass in 25kg (55lb)
The Asian water monitor (Varanus salvator) is second-biggest lizard in the world after Komodo dragon,[1][12][194] and also the longest lizard in the world. The largest specimens come from Malaysia, particularly around the Cameron Highlands, where specimens over 2.5m (8.2ft) in TL (total length) are sometimes encountered. Jasmi (1988) records that wild specimens can weigh up to 25kg (55lb).[4][5] The largest member this species from Sri-Lanka and attained in length of about 3.21m (10.5ft).[1] Thailand is also home to some huge water monitors but, elsewhere the water monitor grows smaller sizes. The biggest found in Java are around 2.1m (6.9ft) of TL (total length), 2m (6.6ft) in Sri Lanka, 1.7m (5.6ft) in Sumatra and mainland India and less than 1.5m (4.91ft) on the island of Flores.[5] Unverified weights of 50, 70 and even 90kg (110, 150 and even 198 pounds) have been reported in escaped pets in Florida but such enormous weights remain unverified.[1] The 80 males slaughtered for the skin trade in Sumatra had an average weight of only 3.42kg (7.5lb), with a snout-vent length of 56.6cm (22.3in) and a total length of 142cm (4.66ft); while 42 females had an average weight of 3.52kg (7.8lb), with a nose-to-cloaca length of 59cm (23in) and a total length of 149.6cm (4.91ft).[195] Among these monitor lizards, some specimens weighed from 16–20kg (35–44lb). Another study in Sumatra by the same authors also estimates the weight of some specimens at 20kg (44lb)[196] while the average adult weight in the population is estimated to be about 7.6kg (17lb).[197]
Crocodile monitor (Varanus salvadorii)
The crocodile monitor considers the longest lizard in the world and third-hugest lizard after Komodo dragon and Asian water monitor
The crocodile monitor (Varanus salvadorii) usually counts the longest lizard in the world.[12] Approved what the biggest specimens can exceed in the length of 3 meters (10ft).[198] The tail of the species is proportionally very long with some sources claiming it to average 210% of the animal’s snout to vent length.[199] According to (Pianka, 2004) the average total length of the lizard ranges between 1,160 and 2,550mm (3.81 and 8.37ft) with the average weight ranging between 5 and 6.38kg (11.0 and 14.1lb).[7] The book, “The Guinness Book of Animal Facts and Feats” cites an individual with a recorded total length of 3.23m (10.6ft)[198] however, such a large size has not been verified.[1] The species can obtain a weight of 20kg (44lb).[200] The largest members of this large species of monitor attained the length in 244cm (8.01ft)[6] and 265cm (8.69ft).[7] Some information suggested what crocodile monitors may measure 5m (16ft) long and 90kg (200lb) mass,[201] although, this has not yet been confirmed.[198]
Nile monitor (Varanus niloticus)
The Nile monitor is the largest lizard native to Africa with a maximum reliable length of more than 2.43m (8.0ft) and a mass in almost 20kg (44lb)
The Nile monitor (Varanus niloticus) is the largest lizard native to Africa alive today and fourth or fifth-largest lizard in the world after Komodo dragon, Asian water monitor, crocodile monitor and competes with perentie.[1] In most cases, males of Nile monitors reach a total length of 150-170cm (4.9-5.5ft) and weigh in the region of 5–10kg (11–22lb),[202] while females are about 1.34m (4.4ft) long and weigh about 3kg (6.6lb). Exceptionally large wild specimens of the Nile monitors can probably reach about 2.5 meters (8.2 feet) in length and weigh about 20kg (44lb), making them one of the largest lizards in the world fauna.[1] In 1929, a photograph of a Nile monitor from South Africa was published, which was said to have reached a total length of 2.13m (7.0ft) and weighed almost 19kg (42lb).[203] The same author also reports that he shot another monitor lizard approximately 2.5m (8.2ft) long.[202] The largest animal this species reliably known to date reached a total length of 242cm (7.94ft) with a length from the muzzle to cloaca length 98cm (39in) and estimated at 12–15kg (26–33lb)[204] According to Faust (2001), the largest representative this species measured 2.43m (7.97ft).[1] A specimen with a length of 188cm (6.17ft) (length from tip of muzzle to cloaca is 74cm (29in)) was recorded from Orange Province in South Africa. Near Lake Chad, the maximum recorded size for a male is 2.07m (6.8ft) and for a female is 1.55m (5.1ft). The largest female Nile monitor for which there is reliable data reached 1.7m (5.6ft). Thus, it seems likely that in South Africa some specimens can reach lengths of 200cm (6.6ft) or more, but they are very rare.[202]
Perentie (Varanus giganteus)
The perentie is the largest lizard Australia, and also one of the biggest lizards in the world
The perentie (Varanus giganteus) is by average length and weight the largest extant lizard native to Australia and fifth or fourth-biggest lizard in the world after Komodo dragon, Asian water monitor, crocodile monitor and competes with Nile monitor.[1] Its endemic to arid central Australia, found west of the Great Dividing Range. The largest individual recorded by (Stokes, 1846) measuring 2438mm (8ft) in total length.[1] However based on the vagueness of this account some authors claim Stokes’ reported measurement of 2438mm (8ft) is exaggerated and a more likely maximum length may instead be 2.3m (7.5ft) (Stirling, 1912).[1] There have been larger individuals supposedly measuring more than 2.5m (8.2ft) in total length and 20kg (44lb) in mass reported but such claims remain unverified.[1] The maximum size of the perentie is probably not as great as many authors claim King & Green (1993) provide a useful discussion of size in this species. The longest found on Barrow Island by King et al. (1989) had a total length of 1.96m (6.4ft) with a SVL in 88cm (35in) and the heaviest weighed 11.7kg (26lb). Butler (1970) record s a specimen of 17kg (37lb) with a total length of 1.93m (6.3ft) from the same island.[1][205] Stokes (1846) records that two specimens collected on Barrow Island in 1840 had total lengths of 2.13m (7.0ft). Strimple (1988) suggested that one of these animals was the type specimen used by Gray, which has a total length of only 2.02m (6.6ft). Many perenties do not grow to such an enormous size, and the specimens on Barrow Island may grow larger than the main population. The limited data available suggests that females reach a smaller maximum size than males, rarely exceeding 1.4m (4.6ft) TL. Bredl (1987) records males of 1.8 and 1.9m (5ft 11in and 6ft 3in) TL, and a female of 1.3m (4.3ft) TL. It appears that perenties approaching 2m (6.6ft) in length are the exception rather than the rule, and that in general few specimens grow larger than 1.5m (4.91ft). If specimens of 2.4m (7.9ft) or more have ever existed, none seem to have survived to the present.[205][206]
The black-throated monitor (Varanus albigularis microstictus) is a subspecies of rock monitor (Varanus albigularis) and, perhaps, the heaviest lizard in Africa. Although, little information is available on the size of this species of monitor lizards. According to many information black-throated monitor attains 2.1m (6.89ft) in length and 27kg (60lb) in mass.[8][207][208] However, unknown whether this mass is reached in captivity or in the wild. By another – black-throated monitor has a mass of 23kg (51lb).[209] However, specimens 2.13m (7.0ft) for this species rock monitor was recorded.[206][210]
Lace monitor (Varanus varius)
The lace monitor – the second-largest lizard native to Australia and seven-largest lizard in the world
The lace monitor (Varanus varius) is the second-largest lizard in Australia after perentie[9] and seven-largest lizard in the world overall.[12] The largest instances may measuring in the length of 2m (6.6ft) with a SVL of 76.5cm (30.1in) and a mass of 14kg (31lb).[4] In 1986 Krefft suggested that lace goannas grow as large as 250cm (8.2ft).[9] None of this size exist now and specimens of 2m (6.6ft) TL are exceptional. A lizard from Mallacoota, Victoria measured 75cm (30in) SVL, 192cm (6.30ft) TL and weighed 14kg (31lb).[9] Another from Healesville was 192cm (6.30ft) long and weighed almost 20.5kg (45lb).[9] It was found to have eaten four foxcubs, three young rabbits and three large blue-tongued skinks,[9] and all this was taken into account when weighing, is unknown. And, accordingly, the most commonly used maximum size for this species is a length of 192cm (6.30ft) and a weight of 14kg (31lb) for the previously mentioned specimen.[4][7][12] In conu'ast, a large male in the spring measured 1,5m (4.91ft) TL and weighed 4.2kg (9.3lb). Males grow larger than females and probably have larger home ranges.[9]
In the genus Varanus has more 80 species of lizards and all of to 11 subgenera, members which vary considerably in sizes:[4]
Subgenus Empagusia
Bengal monitor is the second-largest lizard in Asia after Asian water monitor
The largest representative of the subgenus Empagusia is Bengal monitor (Varanus bengalensis) with a length in 1.75m (5.7ft) and a SVL of 75cm (30in), a mass of 7.2kg (16lb),[211] and in captivity even more – 10.2kg (22lb).[212] However, in average much less – 1,5m (4.91ft) in a length and 2.7kg (6.0lb) in a mass.[7][213] Even so, by some reports it grows much larger.[7][213][206]
Another large species of this group are the clouded monitor (Varanus nebulosus) with a length of 1.5m (4.91ft) and a SVL of 60cm (24in),[214] the roughneck monitor (Varanus rudicollis), which has a total length of 1.46m (4.8ft), a SVL of 59cm (23in) with a mass of 4kg (8.8lb),[206][215] the Dumeril's monitor (Varanus dumerilii) (Maximum size is about 135cm (4.43ft) TL. A breeding pair maintained in captivity measured 130cm (4.3ft) TL, 2,950g (6.50lb) (male) and 100cm (3.3ft), 2,300g (5.1lb) (female). Wild animals of 30cm (12in) SVL weigh about 1kg (2.2lb))[7][206][216] and the yellow monitor (Varanus flavescens) with a maximum length of 95.2cm (3.12ft), a SVL of 44.8cm (17.6in)[4] and a mass of 1,040g (2.29lb).[206][217]
Subgenus Euprepiosaurus
The largest members of this subgenus are the mangrove monitor (Varanus indicus) and its close relatives with a total length of 1.5m (4.91ft)[4][206][218] (The largest Mariana monitors (Varanus tsukamotoi) found on Guam were 58cm (23in) for male and 44cm (17in) for female in SVL with wrights of 1,900g (4.2lb) and 500g (1.1lb) in weight respectively),[7][206][218] the Rainer Günther's monitor (Varanus rainerguentheri) with a maximum total length of 1.33m (4.4ft), probably even 1.5m (4.91ft),[219][4] the tricolor monitor (Varanus yuwonoi) having the maximum length of 1.46m (4.8ft),[219] the blue-tailed monitor (Varanus doreanus) exceeding 1.35m (4.4ft) long,[7] the Rennell Island monitor (Varanus juxtindicus) reaching 1.33m (4.4ft) in length[7] and the New Ireland monitor (Varanus douarrha) growing the length in 1.33m (4.4ft).[4]
Other large lizards in this group include the quince monitor (Varanus melinus) (maximum length of 1.28m (4.2ft)),[220] the peach-throated monitor (Varanus jobiensis) (a total length of 1.2m (3.9ft) and a SVL of 45cm (18in)),[221] the Mussau Island blue-tailed monitor (Varanus semotus) (a total length of 1.2m (3.9ft) and a SVL of 48cm (19in))[4] and the sago monitor (Varanus obor) (a total length of 1.13m (3.7ft) and a SVL of 44cm (17in)).[4]
Subgenus Hapturosaurus
The largest tree monitor is the blue tree monitor (Varanus macraei) with a maximum total length about 1.1m (3.6ft) and a SVL of 36cm (14in).[7][4] Another large tree monitors are the emerald tree monitor (Varanus prasinus), which can attains 1m (3.3ft) in a total length, 30cm (12in) in a snout-vent length,[222] the black tree monitor (Varanus beccarii) with a maximum size of 34cm (13in) SVL, 94.5cm (3.10ft) total length[223] and the Bogert's monitor (Varanus bogerti) measuring a length in 94cm (3.08ft) with a SVL of 32cm (13in).[4]
Subgenus Odatria
Whilst, in this subgenus have a lot of species, they are the smallest monitor lizards.[4] In its include Dampier Peninsula monitor (Varanus sparnus) – the smallest monitor with a length of 227mm (8.9in), a SVL of 116mm (4.6in) and a mass of 16.3g (0.57oz).[4]
The largest species in this subgenus is the black-palmed rock monitor (Varanus glebopalma) measuring the maximum size of 397mm (15.6in) and 1.1m (3.6ft) for SVL and total length respectively (an average mass constitute 240g (8.5oz) with a SVL of 290mm (11in), for the max.size much higher).[7] Another big member this group is Mitchell's water monitor (Varanus mitchelli), which may grow a length in 60–96cm (1.97–3.15ft)[224] with a SVL in 22–32cm (8.7–12.6in).[7]
Subgenus Papusaurus
The only species Papusaurus is a crocodile monitor (Varanus salvadorii) (see higher).
Subgenus Philippinosaurus
The Gray's monitor along with its relatives is one of heaviest lizards in the world
This are large frugivorous monitor lizards (175cm (5.74ft) long or more with a mass in >8kg (18lb)).[21] The Northern Sierra Madre forest monitor (Varanus bitatawa) with a maximum size instead be 180cm (5.9ft) in length and 9kg (20lb) in mass, the Gray's monitor (Varanus olivaceus), the biggest individual of which was just 188cm (6.17ft) long and a mass of 8.9kg (20lb) and the Panay monitor (Varanus mabitang) with the largest specimen which had a length of 175cm (5.74ft) and a mass in 8kg (18lb),[21][22] are one of the biggest lizards in the world. They measuring of 76.6cm (30.2in), 73cm (29in) and 70cm (28in) SVL respectively.[22]
Subgenus Polydaedalus
The largest species Polydaedalus are the nile monitor (Varanus niloticus) and the black-throated monitor (Varanus albigularis microstictus) (see higher).
Another large member of African monitors is a rock monitor, and to be more precise, its type species – white-throated monitor (Varanus albigularis albigularis) with a maximum length of 2m (6.6ft) and 15–17kg (33–37lb)[225] and an average size of 0.8–1.5m (2.6–4.9ft) in length and 3.2–5kg (7.1–11.0lb) for females and 6–8kg (13–18lb) formales in mass[226][227][228] by some sources. But, however, according to others even mass in 7.7kg (17lb) is a very rarely.[206][210]
Although, large sizes may reach also other members of Polydaedalus. This include savannah monitor (Varanus exanthematicus) and Yemen monitor (Varanus yemenensis) with a length of 100cm (3.3ft) (50cm (20in) SVL)[4][206][229] and 115cm (3.77ft) (59cm (23in) SVL)[4] respectively (First one can weighing don't less 1kg (2.2lb))[229] However, Bosc's monitor in captivity can be extremely obese and exceed 150cm (4.9ft) (75cm (30in) SVL)[4] long and 6kg (13lb)) weight.[229]
The ornate monitor can attains the length of 2m (6.6ft),[4] however, the ornate monitor is not currently considered a separate species, but is merely a polymorphisms of two different species.[230]
Subgenus Psammosaurus
The Caspian monitor is a largest lizard in Central Asia, with a maximum reliable length of 1.4m (4.6ft) and a mass of almost 3kg (6.6lb)
The subgenus Psammosaurus includes in its a large lizards and medium by size monitor lizards, growing the length of don't less 120cm (3.9ft).[4] The hugest member this little group and also, the largest lizard in Central Asia – desert monitor (Varanus griseus) specifically, Caspian monitor (Varanus griseus caspius) with a max.total length of 1.4m (4.6ft), a SVL of 58.5cm (23.0in) and a mass of 2,850g (6.28lb) and a SVL of 46cm (18in) with a mass in 2,700g (6.0lb) for longest and heaviest males and females respectively.[206][231] The Nesterov’s desert monitor (Varanus nesterovi) also quite large monitor attaining 1.2m (3.9ft) in a length and 50cm (20in) in a SVL.[4]
Subgenus Solomonosaurus
The Solomon Island spiny monitor (Varanus spinulosus) is a species of subgenus Solomonosaurus, with a maximum reliable length and mass in captivity of 1m (3.3ft) (SVL 31cm (12in)) and 841g (29.7oz) respectively.[7]
Subgenus Soterosaurus
The largest Soterosaurus is a Asian water monitor (Varanus salvator) (see higher).
Second hugest after Asian water monitor are the marbled water monitor (Varanus marmoratus) and the Palawan water monitor (Varanus palawanensis) with a total length of almost 2m (6.6ft) (SVL of last one measuring 78.8cm (31.0in)).[4][232][233] By the data marbled water monitor can weighing about 7.2kg (16lb),[7] although, since V. palawanensis was previously considered a subspecies of V. marmoratus, which species this weight referred to is unknown.[233]
Even medium-sized monitors this subgenus are quite large. For example, the yellow-headed water monitor (Varanus cumingi) and the large-scaled water monitor (Varanus nuchalis) with maximum length of 1.5m (4.9ft) and 1.45m (4.8ft), a SVL of 60cm (24in) and 53cm (21in) and 2.5kg (5.5lb) and 2.8kg (6.2lb) in max.mass respectively.[4][7] Also they include the Togian water monitor (Varanus togianus) (1.5m (4.9ft) in TL and 65cm (26in) in SVL), the Samar water monitor (Varanus samarensis) (1.4m (4.6ft) long with 53cm (21in) SVL), the Rasmussen's water monitor (Varanus rasmusseni) (1.21m (4.0ft) in length and SVL 46.6cm (18.3in)) and the Enteng's monitor lizard (Varanus dalubhasa) (1.16m (3.8ft) long and 50.5cm (19.9in) in SVL).[4]
Subgenus Varanus
The Argus monitor is third largest Australian lizard with mass more 7kg (15lb)
The largest members of this group are the Komodo dragon (Varanus komodoensis), perentie (Varanus giganteus) and lace monitor (Varanus varius) (see higher).
After their, next place occupies the yellow-spotted monitor or Argus monitor (Varanus panoptes), which also is third biggest lizard in Australia, with a maximum length of 1.6m (5.2ft) (SVL of 74cm (29in)) and mass of 7kg (15lb),[4][7] although in captivity huge individuals can weigh about 10kg (22lb) and even more.[234]
A close relative of the aforementioned species is Gould's monitor (Varanus gouldii), also attains large sizes — exclusively large specimens measuring length of 1.55m (5.1ft) (SVL of 67cm (26in)) with a mass of 5kg (11lb).[206][235] The Rosenberg's monitor (Varanus rosenbergi) grows 1.5m (4.9ft) long,[206][236] probably 1.6m (5.2ft) with a SVL of 70cm (28in).[4]
Albeit, not as long as some other species of large varanids, the Spencer's monitor (Varanus spenceri) can weigh a very large for its length due to its massive and bulky body, and has a length of 1.2m (3.9ft) (SVL of 55cm (22in)) and mass of from 2.25kg (5.0lb)[206][237] till 6kg (13lb)[4][7] depending of sources.
The smallest representative of this genus is the Mertens' water monitor (Varanus mertensi) with a length of 1.3m (4.3ft) (SVL of 48cm (19in)).[4] Though, not as bulky as some other species of monitor lizards, in captivity, it can weigh up to 4.9kg (11lb).[206][238]
Night lizards (Xantusidae)
See also: Xantusidae
The largest night lizard is the yellow-spotted tropical night lizard (Lepidophylum flavimaculatum) which attains a snout-to-vent length (SVL) of 12.69cm (5.00in) with a tail 1/3 times exceeding the body length.
Knob-scaled lizards (Xenosauridae)
The largest xenosaurid is a knob-scaled lizard (Xenosaurus grandis) with a snout-vent length of 12.9cm (5.1in) and a total length of 24cm (9.4in).[239]
See also
Megalania – the largest land lizard to have ever lived
Mosasauridae, an extinct family in the order Squamata which includes the largest lizards of the world
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